Abul-Qasim Babur Mirza was in possession of Jurjan and Mazandaran of Iran as well as Quchan in Khurasan in 1448 C.E. His brother Ala-ud-Daulah Mirza after his defeat at the Battle of Tarnab sought refuge with him at Quchan. Babur Mirza had been preparing for an invasion of Khurasan well before the invasion of Ulugh Beg in that region. He waited till winter of 1448 to make his move. Babur Mirza set out with his army towards Mashad which was abandoned by Abdal-Latif Mirza on the approach of his army and taken. From there Babur marched towards Sarakhs.
Meanwhile, circumstances in Samarkand became serious while Ulugh Beg was away. The leader of the Uzbeks, Abul-Khayr Khan, had invaded Transoxiana and raided the suburbs of Samarkand. Ulugh Beg was at that time chasing after Ala-ud-Daulah Mirza and had reached Bistam and Pul-e-Ibrahim. With this invasion Ulugh Beg had no time to consolidate his control over Khurasan and he left for his beloved city of Samarkand with the body of Shahrukh Mirza leaving Abdal-Latif Mirza in charge of Herat.
On arrival at Sarakhs, Abul-Qasim Babur Mirza sent forces in two directions; one under the command of Khalil Hendugha in direction of Amu Darya to harass Ulugh Beg's forces and the other in direction of Herat. Hinduka attacked the baggage train of Ulugh Beg and captured Ibrahim, the son of Edigu-Timur (an Amir of Ulugh Beg). When Ulugh Beg was crossing the river Amu Darya, he was attacked and looted by the Uzbeks. He then rushed towards the city of Bukhara and stayed the winter there. Meanwhile, when the other forces of Abul-Qasim Babur Mirza moved towards Herat, Abdal-Latif Mirza abandoned the city and was moving towards Samarkand when he was stopped by his father who ordered him to take his position in Balkh.
After Abdal-Latif Mirza abandoned Herat, Yar Ali besieged the city and took it after three days. Finally, after 20 days, the army of Abul-Qasim Babur Mirza arrived and besieged the city yet again. Yar Ali was defeated and captured. He was immediately executed in the bazar of the city town center in February, 1449. The city of Tūn was given over to Ala-ud-Daulah Mirza, but out of fear, jealousy or both, he had him arrested along with his son Ibrahim Mirza and imprisoned in Herat.
The conquest of Khurasan in 1449 by Abul-Qasim Babur Mirza was decisive. Although, it would be briefly taken by Sultan Muhammad Mirza another brother of Abul-Qasim Babur Mirza from Fars in 1450; yet it remained in Abul-Qasim Babur Mirza's hands till his death in 1457.
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The original article can be found at Babur Mirza's Invasion of Khorasan and the edit history here.